Alan - sorry to read of your losses.
See our PG 2 with regards how to register for the first time where the deeds have been lost/destroyed -
First registration of title if deeds are lost or destroyed (PG2) - GOV.UK
With regards the deaths and how to then deal with the property see our online assisted guidance -
HMLR Guide Page · HM Land Registry
I have linked you to the guidance where there were joint owners and then a sole surviving owner after the first death. The executor for your late Mother would be the one to then deal with the property re a sale. However as the deeds are lost and there is no registered title any buyer is very likely to insist that it's registered first. Registering it in the name(s) of the beneficiary is the usual way to go in such cases.
Any first registration will therefore include a copy of your Father's death certificate and Mother's probate